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N1 - Funding Information:This research was supported through NIH ( R01HL126845 ) and Muscular Dystrophy Association ( MDA514335 ) grants to A.K., NIH ( R01HL134824 ; R35HL135754 ) to P.J.M., NIH ( P41-GM104601 , R01-GM122420 , and R01-GM123455 ) to E.T., and NIH ( R01AR045653 and R01HL045565 ) to T.A.C. C.M. was supported by American Heart Association post-doctoral fellowship ( 16POST29950018 ). S.B. was supported by the NIH Tissue microenvironment training program ( T32-EB019944 ). J.H. was supported by Beckman-Brown postdoctoral fellowship. All simulations were performed using the Blue Waters PRAC allocation (grant ACI1713784 to E.T.) of the National Center for Supercomputing Applications at UIUC . Three cores at UIUC supported this project: transgenic mouse facility core, high-throughput sequencing and genotyping core, and histology-microscopy core. Cardiovascular phenotyping was carried out in part in the Biomedical Imaging Center at the Beckman Institute for Advanced Science and Technology at the University of Illinois at Urbana-Champaign and in the Mouse Cardiovascular Phenotyping Core at the Washington University School of Medicine , St. Louis, MO.Funding Information:This research was supported through NIH (R01HL126845) and Muscular Dystrophy Association (MDA514335) grants to A.K. NIH (R01HL134824; R35HL135754) to P.J.M. NIH (P41-GM104601, R01-GM122420, and R01-GM123455) to E.T. and NIH (R01AR045653 and R01HL045565) to T.A.C. C.M. was supported by American Heart Association post-doctoral fellowship (16POST29950018). S.B. was supported by the NIH Tissue microenvironment training program (T32-EB019944). J.H. was supported by Beckman-Brown postdoctoral fellowship. All simulations were performed using the Blue Waters PRAC allocation (grant ACI1713784 to E.T.) of the National Center for Supercomputing Applications at UIUC. Three cores at UIUC supported this project: transgenic mouse facility core, high-throughput sequencing and genotyping core, and histology-microscopy core. Cardiovascular phenotyping was carried out in part in the Biomedical Imaging Center at the Beckman Institute for Advanced Science and Technology at the University of Illinois at Urbana-Champaign and in the Mouse Cardiovascular Phenotyping Core at the Washington University School of Medicine, St. Louis, MO. C.M. S.B. and A.K. conceived the project and designed the experiments. C.M. S.B. F.L. K.L. D.J.P. S.N.K. E.R.L. N.P.M. J.H. and L.W.D. performed experiments. T.A.C. and P.J.M. provided reagents. C.M. S.B. K.L. S.N.K. E.R.L. N.P.M. E.T. P.J.M. and A.K. interpreted results and wrote the manuscript. All authors discussed the results and edited the manuscript. The authors declare no competing interests.

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